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An epub is basically a zipped folder of htm,css,images etc. and opf,ncx etc.. Some have sub-folders containing the various elements.
When using Sigil it automagically creates the correct sub-folder structure (inside the epub) and moves all parts to their correct paths, and updates the necessary paths in those files, so it becomes a fully compliant correct epub (most work anyway, but ignore the correct sub-folder structure).
It's this internal structure we are talking about when referring to Styles folder etc., thus when a (x)html file is using any css or image file thats why it needs the ../ to navigate to the root folder then the folder required 'Styles' or 'Images', so a link reference would then become '../Styles/filename.css' or '../Images/filename.jpg'
Also as said by other this link reference is case-sensitive.
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