I offer a wild speculation. From looking a little bit into libnickel during the last days, I have got the impression, that there line-height is primarily set in em. Even if readingLineHeight=1.3 seems to suggest a unit-less value, it might perhaps be handled internally as 1.3em. This difference between the internal line-height setting in em and your css setting might produce differences in elements that inherit the line-height.
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