I bought my Sony 505 from Waterstones. The person I spoke to there seemed pretty knowledgable and was helpful. Waterstones do seem to have the Sony readers on display still so that you can play with them.
I would strongly suggest that you do try to get your hands on any reader you like the look of on paper before you buy. Many people dislike certain readers which others think are great and vice versa. There's no substitute for a hands-on try to assess your own personal preference.
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