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Originally Posted by nrapallo
Not so, since any split file doesn't require an anchor to go to the top of that new file i.e where the 300K split took place. You would only effectively need the anchor links when dealing with illustrations or sub-headings which require linking to a place not at the top of a page or beginning of a file. 
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Which is basically the scenario I was referring to - and level 2-3 TOC elements often falls under this category.
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True, and given the frequency that these secondary links may be used, leaving them in should not be a problem. 
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No, it isn't usually isn't a problem - just so long as the book isn't too large.
An example of a book that is very much affected by anchor tags is The Bible. This is a very large book. with a very large 2 level TOC (at least my version is like this anyway...). If anchors are present in the toc.ncx file, even page turns can get very slow at the end of the book (the New Testament for example). And this is true even when anchors are only present for the secondary TOC elements.
In this situation, the best solution I found was to be very generous with page breaks in the source document/calibre chapter detection. Having a page break at every primary and secondary TOC element eliminated the need for any anchors, and the resulting epub was almost as fast to navigate on the 505 as an LRF. The downside to this is the length of time it takes to do the conversion through calibre - it takes a while to split a file into 1,190 separate files.