Quote:
Originally Posted by Quoth
Mostly they are.
Thunderbird
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Thunderbird is an odd duck, if you'll pardon the pun. Early versions use flat mbox files for each "folder", but some <elided> chose to completely rewrite the backing store to use a homebrew database system on top of the flat mbox files. JWZ has ranted about how stupid this design is so I don't need to elaborate. More recently, Thunderbird can use a simplified Maildir format instead. Maildir as the name suggests is a directory structure for mail messages, where each mail folder is a directory and each message is a single file.
I use Claws-Mail which has always, to my knowledge, used Maildir-like hierarchies for local mail storage and for caching IMAP messages.