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There are a few errors that don't go to the actual source of the error.
Even epubCheck shows the same behaviour at times. One that comes to mind is forgetting to add properties="svg" in the opf. epubCheck will tell you it is missing, but not take you to where it should be fixed. It won't even open the file.
As for the publisher... some do weird things.
Also, I'd rethink not linking the stylesheet. Calibre is quite forgiving with errors, ereaders may not be. For a 6 second job, I wouldn't even question it. I would just do it.
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