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That's the conclusion I came up with as well. So it must come down to being converted to a kindlebook that's causing the issue. I would simplify the ccs as much as possible before converting. If it converts and displays properly, "complicate it" little by little until it displays improperly after conversion. Then you'll have your culprit.
Also consider reading the Kindle publishing guidelines in order to determine if everything you're doing in your epub is even supported for kindlebooks.
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