The open tool drop down will show (and open) the last dozen or so books accessed - but it doesn't show their locations.
And I can't think of any indirect way - e.g. by opening the book in the editor. it will use a working copy of the unpacked epub in calibre's temp location.
It maybe available via the viewers annotation feature, if they can be stored as sidecar files, maybe someone who uses it could comment.
BR
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