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if the original intended for paper is mostly 2 page spreads, then assume person reads in landscape and combine each pair of source images as single landscape images with width: 95% (or 100%) and height: auto.
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Even when I'm using apple's Books in landscape mode, even with each image with width:100% height:auto margin:0, it still displays a gap between the images, but this doesn't happen with other books I have.
