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So I was slightly wrong about the first situation; if using "Update EPUB Always", the cover doesn't change (regardless of the "Update EPUB Cover?" value), so you have to update from metadata to see it change even with "Update EPUB Cover?: Disabled".
I'd assume the same applies to "Update EPUB if New Chapters" in the case of there actually being new chapters, but I have to use "Always" because I can't really count on chapter updates while testing.
Using "Overwrite Always" resets everything back to the original state, with no "cover_image" metadata for the fics that don't have a cover or first image, or a value of "first" instead of "old".
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