Hi Enigma90,
I also have a Glo HD. Last fall I installed firmware 4.25.15875 specifically to get that dictionary because the one in the original firmware was less than adequate. Yes, there were some slight interface changes, but from here I can't tell you what they were.
One other great benefit is the ability to add additional custom dictionaries easily. I added the Websters1913, and the Chambers dictionaries, along with some of the languages.
This has made my use of the reader so much more pleasant. In fact when I read on my old Kindle keyboard model, currently a Proust biography, I curse my self for putting it on that reader, and have reached for the dictionary a hundred times, before I remembered.
I do remember when that firmware came out last fall there was some questions along the line of yours, but nothing seemed to come of it. I have not seen any speed or other usage changes.
I'm sure someone who really knows will be along and give you a more definitive answer, I'll be watching your thread here to see as well.
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