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Interesting, I just tried a few conversions.
Take an epub that's left-aligned, convert to make it justified.
View the original_epub, and it has the original left-aligned text.
Convert it back to left-aligned with the original_epub in place.
View the original_epub, and it is still left-aligned. If Calibre made a new original_epub on the second conversion, that should be justified.
So it seems on conversion, if an original_epub is present, it is just ignored. Not restored, as OP suggests, but just left alone.
Seems strange; if you polish a book with an original_epub, you are warned that it will work on the original and over-write the epub itself. Converting has different behaviour.
This means that if you are converting and want that back-up, you had better delete the original_epub each time.
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