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It could also mean some difference in the screen configuration.
For instance, the librie and the prs500 are pretty technically identical. Case in point, you can read files for either one on either one if you don't mind about slight glitches in the formatting: because of the permanent status bar that's part of the prs500 display at the bottom, its screen's real estate is diminished as compared to that of the librie.
That means that a lrf file meant for the prs500 will leave a blank at the bottom of each pages on the librie, and on the prs500 a file meant for the librie will actually display each pages (as they'd be originally) over two.
I tend to be very cynical about such things, but it could be something as simple as that.
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