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Calibres cover exists only in Calibre. What cover, depends how it was obtained.
It could be extracted from a the book (in the MDE or part of the import)
It could be created by a Plugin Builtin or (User supplied)
Downloaded with Metadata
Dropped into the Comments or MDE from any external (to calibre) source
There are a number of Actions that cause the Library Copy of the book to get updated with the current.
Sending to a Device will only update the SENT copy and my modify the dimensions based upon the selected profile
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