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The original filename is only used to import metadata. Once the book is in calibre, calibe creates files/folders that look like this:
authors/title (id)/title - authors.ext
Whatever is in those fields in the library GUI will be used to construct the calibre library filename. If you want to see FN LN in the library GUI but LN, FN in the Box.com file view, I would suggest using the Save to Disk toolbar icon to export copies of your books into your local Box.com folder.
See Preferences ==> Import/Export ==> Saving books to disk to configure how they will get saved.
Note: the default should be fine, as it uses:
author_sort/title/title - authors.ext
e.g. using a default setup it should already be in LN, FN
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