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The "overrides" are just class names like any other.
If you don't have a @font-face, there's no way the ebook reader will know which font file to use when you say "Merriweather Regular" (no, even if the font is named exactly like that, readers are not that smart). @font-face rules provide a link between the family name you use in font-family and the actual file embedded in the ePub (and listed in the manifest). Without that, it may work in web pages (because browsers in a computer have access to all fonts installed, and the computer keeps an index of which font name corresponds to which file), but not in an ePub.
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