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Will, I agree with you on the bullet-to-barrel seal but I think most breeches are not atmosphere sealed (vented to at least reduce 'kaboom' damage). Ah, just had a thought though, the breech doesn't matter because the cartridge brass will seal that end of the barrel during the burn. So indeed combustion must be sufficient with just the atmosphere inside a given cartridge.
Which is also interesting because some cartridges are so full of powder versus others, thinking of "compressed" rifle loads versus, say, the small powder load in .357 Magnum. Must not take much oxygen to burn even a bunch of powder.
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