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For eBooks I'd imagine it would be "copies sold" or if someone wanted to boost their numbers, "copies downloaded", especially if there had been a free giveaway promo at some point.
For print books, it's kind of a measure of how many the publisher was able to print out and try and ship to the stores, based on anticipated response and actual sales.
So even the stuff that gets remaindered on the sale table gets counted because a book was physically produced and sent out, even if it just went into storage in a warehouse and ultimately recycled, rather than into a home or library.
So yeah, likely "copies downloaded", since they usually put in-print numbers on the blurbs to make the author/title look like they've been really popular.
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