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The only ebook stores I know of that do require ISBNs for ebooks are the ones you mentioned, Kate - Sony and Apple. None of the other ebookstores that accept independently published ebooks, including the other retailers that Smashwords distributes to (Kobo, Diesel, etc) require ISBNs, and nor do Amazon or B&N. As mr ploppy points out, though, if you use Smashwords to publish to Sony and Apple, you can get an ISBN from Smashwords for little or no cost. The same applies to publishing your ebook via Lulu (for distribution to Apple iBookstore).
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