If all you are doing is checking books out of the library you probably don't need wireless.
If you are someone who is buying books the WiFi or 3G option is wonderfully liberating. You don't have to own a computer to own and use a Kindle to its fullest. I am assuming the same is true for the Nook.
When I travel, I can download books on the fly. I don't have to worry about carrying a USB cord with me and finding a computer to download a book. I don't travel with a laptop or a netbook so having a computer is not a guarentee. Or if I do have it with me and I am in an airport I want to be able to download the book I want without having to pull out my laptop.
Sony's are the most expensive 6 inch e-readers and are the only one of the big four (Sony, Amazon, B&N, and Kobo) to not have a wireless option. I find that reasonably silly on Sony's part.
I know that some people place touchscreen before wireless but I would guess that most of the folks buying e-readers prefer wireless over touch screen. It is easy enough to change a page using a button. It might be more convienent (sp) to use a swipe but I would find it much more inconvienent to have to find a computer to download a book on the fly.
And yes, I have ordered many a book, normally a sequel, from an airport and even on the metro.
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