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Originally Posted by Stinger
Its the metadata for that book. I believe it's created so if you want to import those epubs into a different program, you can do so with all your nice metadata intact.
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It's also used when adding the book to Calibre. If that file exists when you try to import into Calibre, it will override any other metadata in the file or in the filename. If you want to give a book to a friend who's also using Calibre, give them the opf file too.
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At least for epubs, I believe the metadata is also stored within the actual epub
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Yes, it's in the content.opf file inside the EPUB.