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I don't think the Kindle 1 had it (maybe started with the Kindle 2??), but the later Kindle's have support for actual page numbers that match up to the page numbers in a specific paper edition of a book. It's not available on all books as some books are digital only and others just don't have them because Amazon or the publisher haven't mapped the pages out. If a book has them it's description on the Amazon website will usually have a line that says "Page Numbers Source ISBN:"
Some ePub based readers use "page numbers" (as opposed to locations on the Kindle), but in general they don't relate to any paper edition of a book. IIRC the Nook does do "real" page numbers on some books, but I'm not sure.
Some other ePub devices use page numbers that equate to one screens worth of text, therefore the number of "pages" in a book changes any time you adjust font size, margins or line spacing.
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